The once peaceful island of Swarovski was destroyed by the genius invention of Flinder Lockwood (voiced by Bill Hader), the Re-Eater. In the end, he successfully destroyed the Re-Eater and planned to rebuild a beautiful home with Sam (voiced by Anna Faris). At this time, Flinder's most admired idol, Chester V (voiced by Will Forte), the boss of Liv Company, fell from the sky. Chester is a genius inventor, and his company has gathered the world's top scientists. Chester invited Flinder to join Liv Company and promised to take the islanders to California, and his company will be responsible for cleaning up the dilapidated island. In order to become a fantasy inventor, Flinder tried his best to carry out research and development. At the same time, bad news came from his hometown. Many cleaners were captured by living food. It seems that the Re-Eater was not completely destroyed and continued to develop in a direction that Flinder had never expected. Under Chester's order, he and Sam, Dad and other partners embarked on a journey back home to find a solution to the trouble...