Since the birth of the earth, a strange creature has appeared on this blue planet. They have been evolving and transforming, and finally became the familiar appearance of the Minions. The Minions have spent their lives looking for the evil boss and are proud to be his subordinates, but millions of years of efforts have finally turned into nothing. After a long period of silence, the Minions named Kevin decided to go on the road again. He brought Stuart, who loves music, and Bob, who is more likely to fail than to succeed, across the ocean to the United States in 1968. The three little guys accidentally heard that Orlando was about to hold a "Villain Conference", so they hurried over and became the followers of the female devil Scarlett · Killer (voiced by Sandra Bullock). Scarlett soon gave them a task, which was to steal the crown of the Queen of England. The little guys who didn't know the importance of the matter went out with a smile, and what awaited them would be an unprecedented adventure...