The story takes place in Harajuku, a bustling city filled with countless fashion elements. There is a store called "PARK" here. The gentle and considerate Sudo Rito (voiced by Haruna Luna), who is not good at expressing her feelings, the very individual Shirako Mari (voiced by Uesaka Sumire), who has her own opinions and is not easily shaken, and the highly intelligent Wataru Kotoko (voiced by Iwami Goka), who is easily trapped in her own world, run this store together. What the three did not expect was that one day, aliens actually attacked the earth. Not only that, humans were at a disadvantage and almost lost. At this time, a mysterious girl named Maruno Misa (voiced by Amano Koka) appeared in front of the three. She and the three girls who love the Harajuku style very much seem to have become the last hope of mankind.