Fighter pilot Yuichi Kanan (voiced by Ryo Kase) reports to the base, and what awaits him is a brand new living environment and familiar flight combat. Here, Yuichi Kanan meets a variety of comrades-in-arms and the unsmiling commander, Kusanagi Mizuso (voiced by Rinko Kikuchi). Yuichi Kanan gets a new fighter, but his comrades in the base are silent about the former pilot of the fighter. So Yuichi Kanan starts to pay attention to the relevant news, and he gradually learns that the former pilot has passed away, and the incident itself has a great relationship with Kusanagi Mizuso. As the investigation deepens, Yuichi Kanan gradually discovers the shocking truth, and his daily battles with the enemy seem to hide meanings that he does not know.